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466 Biographical Memoirs He entered Cambridge in October 1938 and graduated in 1941 with an upper second. His interests were broad and he clearly spent a great deal of time playing soccer (he was captain of the College team), cricket and tennis.
Alan Robertson 467 Alan’s personal life took a rapid change after joining NABGRO in Hendon. He met Margaret (Meg) Bemheim, who was the secretary there and had previously been in ORS Fighter Command, and they married in January 1947. In April they left for the United States where Alan received some training in genetics and animal breeding. He spent three months with Sewall Wright in Chicago and six months with Jay L. Lush in Ames, who between them had done more than anyone to establish a sound theoretical foundation for animal genetics in relation to animal improvement, and to establish and disseminate animal breeding practices based on these principles.
470 Biographical Memoirs population improvement. With Rendel he showed that longevity, so highly regarded by breeders, was a much less important trait than lactation yield in terms of efficiency (5). Breed structure was investigated and it was shown that a hierarchy of breeders of Friesian cattle was already established, with movement of genes down this pyramid (3, 15, 16). It was therefore not surprising that only a small proportion of the variation between herds was found to be genetic (28), so selection of bulls on the basis of progeny yield but ignoring herd yield would not be efficient.
472 Biographical Memoirs were obtained (49), necessary parameters for the contemporary comparison, and analyses undertaken of genetic variation in non-fat components of milk (43), which at that time were not considered in breeding programmes and did not become incorporated until over a decade later.